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I thank Francisco Ayala and John Avise for organizing this conference, Howard Ochman for critical comments on a draft of the manuscript, Daniela Takiya and Roman Rakitov for use of their photograph, Becky Nankivell for preparation of the manuscript and Fig. 9.1, and two reviewers for helpful comments. The work on symbionts of aphids and sharpshooters was supported by National Science Foundation Awards 0313737 and 0626716.

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