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tological and neontological insights takes on special urgency with the acceleration of extinction rates in the modern world, and the incorporation of the spatial dimension offers a powerful vehicle for that integration.


I thank John Avise, Francisco Ayala, and Stephen Hubbell for inviting me to participate in this colloquium; J. Alroy, J. C. Avise, P. G. Harnik, S. M. Kidwell, M. J. Novacek, and J. W. Valentine for manuscript reviews; K. Roy and J. W. Valentine for many fruitful discussions and collaborations; A. Z. Krug and P. G. Harnik for discussions; and P. G. Harnik for assistance with multiple logistic regression. The National Science Foundation, the National Aeronautics and Space Administration, and the John Simon Guggenheim Foundation supported this research and synthesis.

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