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the following undergraduate and Master degree-level field assistants: Sylvie Moulin, Thomas Stevenson, Betsy Abrams, Sera Young, Brendan Hug, Lindsey Boynton, Joy Rowe, Riley Bove, Anne Lyne, Jean Baptiste, David Plate, Joanna Rogers, Robin Martz, Christine Sauve, and Russell Herrold. I thank Kathy Welch for statistical advice and Claudius Vincenz, Laura Betzig, and two anonymous reviewers for helpful comments. This research was supported by the Louis S. B. Leakey Foundation, National Science Foundation Grants SBR 9727229 and BCS 1029056, the American Philosophical Society, and National Institutes of Health Grant 09-PAF00653.

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