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Manyaas for their help in the field; and J. Avise, S. Emlen, P. Sherman, and especially C. Botero, S.-F. Shen, and D. I. Rubenstein for providing discussion and/or comments on earlier versions of the manuscript. The Kenyan Ministry of Education, Science and Technology, the National Council on Science and Technology, the National Museums of Kenya Ornithology Section, the Kenya Wildlife Service, and the Mpala Research Centre enabled this work. D.R.R.’s long-term research was supported by Columbia University, the Howard Hughes Medical Institute, the Smithsonian Institution, Cornell University, the Miller Institute for Basic Research at the University of California at Berkeley, and the Museum of Vertebrate Zoology, as well as by grants from the National Science Foundation (IBN-407713), the American Museum of Natural History Chapman Fund, the American Ornithologists’ Union, the Wilson Ornithological Society, the Society for Integrative and Comparative Biology, the Animal Behaviour Society, the Harvard Travellers Club, the Society of Sigma Xi, Cornell University, the Cornell Laboratory of Ornithology Benning Fund, and Cornell Sigma Xi.

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